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Song, W. q., X. l. Li & Y. l. Chen. 1989. Studies on the karyotype evolution and relationship of Maloideae. Pp. 327--333 in D. Hong (editor), Plant Chromosome Research 1987.
Ekimova. 2005. New chromosome numbers of relict shrubs species of Rosaceae family. Pages 32--33 in Karyology, Karyosystematics and Molecular Phylogeny. St. Petersburg, Russia.